1945, in World War II Germany, the tough Sergeant Don 'Wardaddy' Collier commands a tank and survives a German attack with his veteran crew composed of Boyd 'Bible' Swan, Trini 'Gordo' Garcia and Grady 'Coon-Ass' Travis. He receives a rookie soldier Norman Ellison as the substitute for his deceased gunner and he tries to harden the youth along the way.